Mammalia - Carnivora - Ursidae

Parent taxon: Ursidae according to J. Chorn and R. S. Hoffman 1978

Sister taxa: Adelpharctos, Amphicynodon, Amphicynodontinae, Dinocyon, Gulonina, Harpaleocyon, Hemicyon, Hemicyoninae, Metarctos, Myadina, Taxina, Thaumastocyon, Tremarctinae, Ursinae, Ursulus

Subtaxa: Agriotheriini

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Ecology: ground dwelling herbivore-carnivore