ammonites
Cryptobranchus guildayi Holman 1977 (hellbender)

Amphibia - Caudata - Cryptobranchidae

Full reference: J. A. Holman. 1977. The Pleistocene (Kansan) herpetofauna of Cumberland Cave, Maryland. Annals of Carnegie Museum 46(11):157-172

Belongs to Cryptobranchus according to R. Estes 1981

Sister taxa: Cryptobranchus alleganiensis, Cryptobranchus saskatchewanensis

Type specimen: CM 20470, a mandible. Its type locality is Cumberland Cave, which is in an Irvingtonian cave horizon in Maryland.

Ecology: