Rhynchonellata - Athyridida - Athyrididae

Full reference: G. A. Cooper and R. E. Grant. 1976. Permian Brachiopods from West Texas, IV. Smithsonian Contributions to Paleobiology 21:1923-2607

Belongs to Cleiothyridina according to P.R. Hoover 1981

Sister taxa: Athyris (Cleiothyridina) bajtuganensis, Athyris (Cleiothyridina) epigona, Athyris (Cleiothyridina) warchensis, Cleiothyridina accola, Cleiothyridina acutomarginalis, Cleiothyridina ailakensis, Cleiothyridina anabathra, Cleiothyridina attenuata, Cleiothyridina bajkurica, Cleiothyridina baracoodensis, Cleiothyridina barbata, Cleiothyridina capillata, Cleiothyridina circularis, Cleiothyridina ciriacksi, Cleiothyridina corculum, Cleiothyridina dalmiriensis, Cleiothyridina davidsoni, Cleiothyridina deroissyi, Cleiothyridina dilimensis, Cleiothyridina echidniformis, Cleiothyridina excavata, Cleiothyridina globulina, Cleiothyridina grossula, Cleiothyridina hayasakai, Cleiothyridina interposita, Cleiothyridina intonsa, Cleiothyridina laminosa, Cleiothyridina laqueata, Cleiothyridina macleayana, Cleiothyridina maynci, Cleiothyridina mulsa, Cleiothyridina nielseni, Cleiothyridina nikolaevi, Cleiothyridina orbicularis, Cleiothyridina ovalis, Cleiothyridina pectinifera, Cleiothyridina perthensis, Cleiothyridina pijaensis, Cleiothyridina pilularis, Cleiothyridina rara, Cleiothyridina rectimarginata, Cleiothyridina saraiensis, Cleiothyridina semiconcava, Cleiothyridina seriata, Cleiothyridina shenshuensis, Cleiothyridina simulans, Cleiothyridina solovjevae, Cleiothyridina subexpansa, Cleiothyridina tribulosa, Cleiothyridina tschironensis, Cleiothyridina uralica, Cleiothyridina xetriformis, Cleiothyridina zhexiensis, Cleiothyridellina accoliformis

Type specimen: USNM 154387a, a shell (Articulated shell)

Ecology: stationary low-level epifaunal suspension feeder

Average measurements (in mm): shell 4.58 x 4.21