Actinopteri - Clupeiformes

Parent taxon: Clupeoidei according to R. Betancur-R et al. 2013

Sister taxa: Chirocentridae, Clupavidae, Clupeidae, Clupeoidea, Dussumieriidae, Pristigasteridae

Subtaxa: Coilinae, Engraulinae

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Ecology: nektonic carnivore