Actinopteri - Clupeiformes

Parent taxon: Clupeiformes according to R. Betancur-R et al. 2013

Sister taxa: Alisea, Anchoa, Austroclupea, Chirocentroidea, Clupeidarum, Clupeoidei, Daitingichthys, Etringus, Etrumeus, Ganoessus, Ganolytes, Gasteroclupea, Hacquetia, Harengula, Ilisha, Opisthonema, Osteoglossoidei, Pomolobus, Pseudoellimma, Pseudohilsa, Quisque, Sardina, Sardinella, Sardinops, Sarmatella, Stolephorus, Xyne

Subtaxa: Denticipitidae, Paleodenticeps

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Ecology: nektonic carnivore