Full reference: G. A. Cooper. 1976. Lower Cambrian brachiopods from the Rift Valley (Isreal and Jordan). Journal of Paleontology 50(2):269-289
Sister taxa: none
Type specimen: HU 35160a, a shell (Ventral valve). Its type locality is Nimra Formation, (southern) Negev, Israel - Cooper 1976, which is in the Lenian of dolomite the Nimra Formation Israel.
Ecology: stationary low-level epifaunal suspension feeder