Full reference: M. Matsukawa, K. Shibata, R. Kukihara, K. Koarai, and M. G. Lockley. 2005. Review of Japanese dinosaur track localities: implications for ichnotaxonomy, paleogeography and stratigraphic correlation. Ichnos 12(3):201-222
Sister taxon: Schizograllator xiaohebaensis
Type specimen: OTK-001-3, a footprint. Its type locality is Kuruma, Otari tracksite, which is in a Pliensbachian delta plain siliciclastic in the Kitamatadani Formation of Japan.
Ecology: ground dwelling carnivore