Full reference: M. Zhu, W. Zhao, L. Jia, J. Lu, T. Qiao and Q. Qu. 2009. The oldest articulated osteichthyan reveals mosaic gnathostome characters. Nature 458:469-474
Sister taxa: none
Type specimen: IVPP V15542.1-6, a skeleton. Its type locality is Kuanti Formation, Qujing County, Eastern Yunnan Province, which is in a Ludlow marine siltstone/shale in the Kuanti Formation of China.