Full reference: J. G. M. Thewissen, E. M. Williams, and S.T. Hussain. 2001. Eocene mammal faunas from northern Indo-Pakistan. Journal of Vertebrate Paleontology 21(2):347-366
Sister taxa: none
Type specimen: H-GSP 97193, a set of teeth (left [upper] M1-2 and several enamel fragments). Its type locality is Gali Jhagir, which is in an Eocene terrestrial mudstone in the Chorgali Formation of Pakistan.
Ecology: ground dwelling browser