Insecta - Coleoptera

Parent taxon: Adephaga according to P. Bouchard et al. 2011

Sister taxa: Amphizoidae, Amphoxyne, Carabidae, Carabiformia, Carabilarva, Chimaerocoleus, Colymbotethidae, Dytiscidae, Eodromeites, Gyrinidae, Haliplidae, Hygrobiidae, Memptus, Necronectulus, Noteridae, Pseudomorphinae, Rhysodidae, Trachypachidae, Triadogyrus, Triaplidae, Tritarsidae

Subtaxa: Charonoscaphinae, Coptoclavinae, Coptoclaviscinae, Hispanoclavinae, Timarchopsinae

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Type: Coptoclavinae

Ecology: