Actinopteri - Pycnodontiformes - Pycnodontidae

Full reference: K. Kumar and R. S. Loyal. 1987. Eocene ichthyofauna from the Subathu Formation, Northwestern Himalaya, India. Journal of the Palaeontological Society of India 32:60-84

Belongs to Pycnodus according to K. Kumar and R. S. Loyal 1987

Sister taxa: Pycnodus flabellatus, Pycnodus granulatus, Pycnodus jugleri, Pycnodus lametae, Pycnodus minutus, Pycnodus orbicularis, Pycnodus platessus, Pycnodus preufsii

Type specimen: VPL/K 593, a tooth. Its type locality is East Babbian Gala Locality, which is in a Ypresian channel lag shale in the Subathu Formation of India.

Ecology: nektonic carnivore