Reptilia - Pterosauria - Pteranodontidae

Alternative combinations: Nyctodactylus gracilis, Pteranodon (Nyctosaurus) gracilis, Pteranodon gracilis

Synonyms: Nyctosaurus bonneri Miller 1971, Pteranodon (Nyctosaurus) bonneri Miller 1971

Full reference: O. C. Marsh. 1876. Notice of a new sub-order of Pterosauria. American Journal of Science 11:507-509

Belongs to Nyctosaurus according to D. M. Unwin 2003

Sister taxa: Nyctosaurus lamegoi, Nyctosaurus nanus

Type specimens:

• Nyctosaurus gracilis: YPM 1178, a set of postcrania (Scapulae, coracoids, dorsal and cervical vertebrae, partial forelimbs). Its type locality is Hackberry, which is in a Santonian/Campanian offshore horizon in the Niobrara Formation of Kansas.
• Nyctosaurus bonneri: SMM 11311, a partial skeleton. Its type locality is near Elkader, which is in a Santonian/Campanian offshore chalk in the Niobrara Formation of Kansas.