Reptilia - Sphenisciformes - Spheniscidae

Full reference: C. P. Tambussi, C. I. Acosta Hospitaleche, M. A. Reguero and S. A. Marenssi. 2006. Late Eocene penguins from West Antarctica: systematics and biostratigraphy. Geological Society, London Special Publication 258:145-161

Parent taxon: Spheniscidae according to C. P. Tambussi et al. 2006

Sister taxa: Anthropodyptes, Anthropornis, Archaeospheniscus, Arthrodytes, Dege, Delphinornis, Duntroonornis, Eretiscus, Eudyptes, Eudyptula, Inguza, Korora, Madrynornis, Marambiornis, Marplesornis, Megadyptes, Mesetaornis, Nucleornis, Pachydyptes, Palaeeudyptes, Palaeeudyptinae, Palaeospheniscinae, Palaeospheniscus, Paraptenodytes, Paraptenodytinae, Platydyptes, Pseudaptenodytes, Pygoscelis, Spheniscinae, Spheniscus, Tereingaornis, Wimanornis

Subtaxa: Tonniornis mesetaensis, Tonniornis minimum

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Type: Tonniornis mesetaensis

Ecology: ground dwelling carnivore