Full reference: L. Doguzhaeva. 2002. Adolescent bactritoid, orthoceroid, ammonoid and coleoid shells from the Upper Carboniferous and Lower Permian of the south Urals. Abhandlungen der Geologischen Bundesanstalt 57:9-55
Sister taxa: none
Type specimen: PIN 3871/301. Its type locality is Sim River, 5 km from Sim along Ufa-Chelyabinsk Road, which is in an Artinskian carbonate limestone in the Russian Federation.
Ecology: fast-moving nektonic carnivore