Mammalia - Theriiformes - Arctocyonidae

Full reference: J. D. Archibald, Y. Zhang, T. Harper and R. L. Cifelli. 2011. Protungulatum, confirmed Cretaceous occurrence of an otherwise Paleocene eutherian (placental?) mammal. Journal of Mammalian Evolution 18:153-161

Belongs to Protungulatum according to J. D. Archibald et al. 2011

Sister taxa: Protungulatum donnae, Protungulatum gorgun, Protungulatum mckeeveri, Protungulatum sloani

Type specimen: OMNH 64985, a tooth (last left upper premolar). Its type locality is Spigot-Bottle locality, which is in a Maastrichtian fluvial-deltaic horizon in the Hell Creek Formation of Montana.

Ecology: ground dwelling omnivore