Full reference: K. E. Mickle, R. Lund, and E. D. Grogan. 2009. Three new palaeoniscoid fishes from the Bear Gulch limestone (Serpukhovian, Mississippian) of Montana (USA) and the relationships of lower actinopterygians. Geodiversitas 31(3):623-668
Sister taxon: Lineagruan judithi
Type specimen: ROM 41809, a skeleton. Its type locality is Bear Gulch (Carnegie Museum Collection), which is in an Arnsbergian open shallow subtidal limestone in the Heath Formation of Montana.