Reptilia - Testudines - Testudinidae

Parent taxon: Testudinidae according to P. Meylan and W. Auffenberg 1986

Sister taxa: Bellia, Cachuga, Damonia, Dipsochelys, Floridemys, Geochelone, Gigantochersina, Hadrianus, Hesperotestudo, Impregnochelys, Indotestudo, Kansuchelys, Kinixys, Miotestudo, Nicoria, Psammobates, Sinohadrianus, Stylemys, Testudininae, Testudo

Subtaxa: Homopus comptoni, Homopus scutella

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Ecology: ground dwelling herbivore-herbivore