Insecta - Coleoptera - Schizophoridae

Full reference: A. G. Ponomarenko. 1969. Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]. Trudy Akademiya Nauk SSSR 125:1-240

Belongs to Hadeocoleus according to A. G. Ponomarenko 1969

Sister taxa: Hadeocoleus gigas, Hadeocoleus pelopius

Type specimen: PIN 2240/172, an exoskeleton. Its type locality is Madygen, PIN collection 2240, Dzhailoucho area, which is in a Carnian lacustrine - small siliciclastic in the Madygen Formation of Kyrgyzstan.

Ecology:

Average measurements (in mm): body length 12.0