Insecta - Diptera - Ceratopogonidae

Full reference: A. R. Palmer. 1957. Miocene arthropods from the Mojave desert, California. United States Geological Survey Professional Paper 294-G:237-280

Belongs to Culicoides according to A. Borkent 2013

Sister taxa: Culicoides abbreviatipennis, Culicoides agamus, Culicoides ambericus, Culicoides annosus, Culicoides antilleanus, Culicoides atratus, Culicoides austerus, Culicoides bicolor, Culicoides bifidus, Culicoides brodzinskyi, Culicoides bullus, Culicoides canadensis, Culicoides carri, Culicoides casei, Culicoides ceranowiczi, Culicoides dasyheleiformis, Culicoides elongatulus, Culicoides eoselficus, Culicoides filipalpis, Culicoides fossilis, Culicoides gedanensis, Culicoides gracilior, Culicoides grandibocus, Culicoides hispanicolus, Culicoides jucundus, Culicoides kaluginae, Culicoides laurae, Culicoides liliputanus, Culicoides mammalicolus, Culicoides mioceneus, Culicoides obesus, Culicoides obuncus, Culicoides prussicus, Culicoides speciosus, Culicoides sphenostylus, Culicoides subgedanensis, Culicoides succineus, Culicoides succivarius, Culicoides tenuipennis, Culicoides truncatus, Culicoides tyrrelli, Culicoides ventralis, Culicoides yoosti

Type specimen: USNM 562003, an exoskeleton. Its type locality is USGS Cenozoic 19057, Mule Canyon, Calico Hills, which is in a Barstovian lacustrine - large shale in the Barstow Formation of California.

Ecology: