Reptilia - Testudines - Trionychidae
Full reference: I. G. Danilov, E. A. Zvonok, E. V. Syromyatnikova and N. I. Udovichenko. 2011. A new species of soft-shelled turtle (Trionychidae) from the Middle Eocene of Ukraine. Proceedings of the Zoological Institute RAS 315(4):399-411
Belongs to Trionyx according to I. G. Danilov et al. 2011
Sister taxa: Trionyx aquitanicus, Trionyx barbarae, Trionyx bowerbanki, Trionyx buiei, Trionyx gangeticus, Trionyx gergensi, Trionyx halophilus, Trionyx henrici, Trionyx hurum, Trionyx incrassatus, Trionyx kansaiensis, Trionyx kyrgyzensis, Trionyx miensis, Trionyx parisiensis, Trionyx phayrei, Trionyx planus, Trionyx rivosus, Trionyx sculptus, Trionyx vittatus
Type specimen: ZIN PH 37/145, a partial skull. Its type locality is Ikovo, which is in a Lutetian marine sandstone in Ukraine.
Ecology: aquatic piscivore-carnivore