Reptilia - Testudines - Emydidae

Full reference: W. G. Joyce, A. Petričević, T. R. Lyson and N. J. Czaplewski. 2012. A new box turtle from the Miocene/Pliocene boundary (latest Hemphillian) of Oklahoma and a refined chronology of box turtle diversification. Journal of Paleontology 86(1):177-190

Belongs to Terrapene according to W. G. Joyce et al. 2012

Sister taxa: Terrapene canaliculata, Terrapene carolina, Terrapene coahuila, Terrapene corneri, Terrapene impensa, Terrapene klauberi, Terrapene marnochi, Terrapene mexicana, Terrapene nelsoni, Terrapene ornata, Terrapene whitneyi

Type specimen: OMNH 58158, a shell (near complete shell). Its type locality is Buckshot Arroyo (OMNH VP locality no. V671), which is in a Hemphillian terrestrial sandstone in the Rexroad Formation of Oklahoma.

Ecology: ground dwelling omnivore