Reptilia - Testudines - Testudinidae

Full reference: R. Lydekker. 1889. Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History) Part III. The Order Chelonia 1-239

Belongs to Testudo according to R. Lydekker 1889

Sister taxa: Testudo atlas, Testudo bosporica, Testudo cautleyi, Testudo cernovi, Testudo cubensis, Testudo escheri, Testudo eurysternum, Testudo gallardoi, Testudo hermanni, Testudo kaiseni, Testudo larteti, Testudo nanus, Testudo praestans, Testudo punjabiensis, Testudo robusta, Testudo spratti, Testudo ulanensis

Type specimen: BMNH R1587, a partial shell

Ecology: ground dwelling herbivore-herbivore