Insecta - Coleoptera - Scarabaeidae

Full reference: J. F. Zhang. 1989. Fossil Insects From Shanwang, Shandong, China 1-459

Belongs to Macronota according to J. F. Zhang 1989

Sister taxa: none

Type specimen: 840121, an exoskeleton. Its type locality is Shanwang, Linqu County, collection 3, which is in a Miocene lacustrine - large mudstone in the Shanwang Formation of China.

Ecology:

Average measurements (in mm): body 14.1 x 5.7