Reptilia - Pelecaniformes - Sulidae

Parent taxon: Sulidae according to J. Cheneval 1993

Sister taxa: Eostega, Masillastega, Microsula, Miosula, Morus, Paleosula, Ramphastosula, Sarmatosula

Subtaxa: Sula avita, Sula clarki, Sula dactylatra, Sula guano, Sula phosphata, Sula pohli, Sula willetti

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Ecology: ground dwelling carnivore