Reptilia - Pelecaniformes - Sulidae

Full reference: H. Howard. 1958. Miocene sulids of southern California. Contributions in Science 25:1-15

Parent taxon: Sulidae according to H. Howard 1958

Sister taxa: Eostega, Masillastega, Microsula, Miosula, Morus, Ramphastosula, Sarmatosula, Sula

Subtaxa: Paleosula stocktoni

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Ecology: ground dwelling carnivore