Insecta - Coleoptera

Parent taxon: Polyphaga according to P. Bouchard et al. 2011

Sister taxa: Angarolarva, Bostrichiformia, Bostrychiformia, Cucujiformia, Derodontiformia, Elateriformia, Eucinetiformia, Geotrupidae, Peltosyne, Plastelater, Staphyliniformia, Tomocoleites

Subtaxa: Scarabaeoidea

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Ecology: