Insecta - Coleoptera

Parent taxon: Scarabaeoidea according to R. A. Crowson 1981

Sister taxa: Aliscarabaeus, Belohinidae, Diphyllostomatidae, Glaphyridae, Glaresidae, Hybosoridae, Hybosorites, Lucanidae, Obitiscarabaeus, Ochodaeidae, Passalidae, Pleocomidae, Scarabaeidae, Septiventeridae, Serrulus, Trogidae

Subtaxa: none

Ecology: