Insecta - Coleoptera

Parent taxon: Scarabaeoidea according to J. F. Lawrence and A. F. Newton 1995

Sister taxa: Acanthoceridae, Aliscarabaeus, Belohinidae, Diphyllostomatidae, Glaphyridae, Glaresidae, Hybosoridae, Hybosorites, Lucanidae, Obitiscarabaeus, Passalidae, Pleocomidae, Scarabaeidae, Septiventeridae, Serrulus, Trogidae

Subtaxa: Chaetocanthinae, Ochodaeinae, Taurocerastinae

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Ecology: