Insecta - Coleoptera

Parent taxon: Scarabaeoidea according to J. F. Lawrence and A. F. Newton 1995

Sister taxa: Acanthoceridae, Aliscarabaeus, Belohinidae, Diphyllostomatidae, Glaphyridae, Glaresidae, Hybosorites, Lucanidae, Obitiscarabaeus, Ochodaeidae, Passalidae, Pleocomidae, Scarabaeidae, Septiventeridae, Serrulus, Trogidae

Subtaxa: Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, Procoilodes, Pulcherhybosorus, Tyrannasorus

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Ecology: