Insecta - Coleoptera - Hydrophilidae

Parent taxon: Hydrophilidae according to J. F. Zhang 1989

Sister taxa: Anacaena, Berosini, Cercyon, Creniphilites, Cretohelophorus, Cretosperchus, Enochrus, Epimetopinae, Georissinae, Helochares, Helophorinae, Horelophinae, Hydrochinae, Hydrochus, Hydrophilinae, Laccobiini, Laccobius, Leptobrochus, Litobrochus, Mesobrochus, Mesydra, Ochthebius, Paladicella, Paraspercheus, Penitus, Philhydrus, Philydrus, Prospercheus, Spercheinae, Sphaeridiinae, Tropisternus, Escheria protogaeae

Subtaxa: Hydrous ebeninus, Hydrous pergrandis

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Ecology: