Full reference: M. G. Lockley, S.Y. Yang, M. Matsukawa, F. Fleming, and S.K. Lim. 1992. The track record of Mesozoic birds: evidence and implications. Philosophical Transactions of the Royal Society of London, Series B: Biological Sciences 336:113-134
Sister taxa: none
Type specimen: KPE 50006, in the Kyungpook National University, Earth. Its type locality is Dukmyeongri Tracks, site 8, which is in an Albian lacustrine carbonate in the Jindong Formation of South Korea.
Ecology: ground dwelling carnivore