Full reference: M. G. Lockley, G. Nadon, and P. J. Currie. 2003. A diverse dinosaur-bird footprint assemblage from the Lance Formation, Upper Cretaceous, eastern Wyoming; implications for ichnotaxonomy. Ichnos 11:229-249
Sister taxa: none
Type specimen: CU-MWC 224.5. Its type locality is Zerbst Ranch Tracksite, which is in a Lancian fluvial sandstone/sandstone in the Lance Formation of Wyoming.
Ecology: ground dwelling carnivore