Reptilia - Pelecaniformes - Sulidae

Full reference: G. Mayr. 2002. A skull of a new pelecaniform bird from the Middle Eocene of Messel, Germany. Acta Palaeontologica Polonica 47(3):507-512

Parent taxon: Sulidae according to G. Mayr 2002

Sister taxa: Eostega, Microsula, Miosula, Morus, Paleosula, Ramphastosula, Sarmatosula, Sula

Subtaxa: Masillastega rectirostris

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Ecology: ground dwelling carnivore