Full reference: R. W. Gess, M. I. Coates, and B. S. Rubidge. 2006. A lamprey from the Devonian period of South Africa. Nature 443:981-984
Sister taxa: none
Type specimen: AM5750, a skeleton. Its type locality is Waterloo Farm, which is in a Famennian paralic horizon in the Witpoort Formation of South Africa.