Reptilia - Sphenisciformes - Spheniscidae
Full reference: A. Myrcha, P. Jadwiszczak, C. P. Tambussi, J. I. Noriega, A. Gazdzicki, A. Tatur, and R. A. Valle. 2002. Taxonomic revision of Eocene Antarctic penguins based on tarsometatarsal morphology. Polish Polar Research 23(1):5-46
Parent taxon: Spheniscidae according to A. Myrcha et al. 2002
Sister taxa: Anthropodyptes, Anthropornis, Archaeospheniscus, Arthrodytes, Dege, Delphinornis, Duntroonornis, Eretiscus, Eudyptes, Eudyptula, Inguza, Korora, Madrynornis, Marplesornis, Megadyptes, Mesetaornis, Nucleornis, Pachydyptes, Palaeeudyptes, Palaeeudyptinae, Palaeospheniscinae, Palaeospheniscus, Paraptenodytes, Paraptenodytinae, Platydyptes, Pseudaptenodytes, Pygoscelis, Spheniscinae, Spheniscus, Tereingaornis, Tonniornis, Wimanornis
Subtaxa: Marambiornis exilis
Type: Marambiornis exilis
Ecology: ground dwelling carnivore