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Ceratopsidae

Taxonomy

Ceratopsidae was named by Marsh (1888). It is not extant. Its type is Ceratops. It was considered monophyletic by Sereno (2000).

It was considered a nomen dubium by Lull (1906).

It was assigned to Dinosauria by Marsh (1888); to Ornithopoda by Mansel-Pleydell (1902); to Orthopoda by Depéret (1896), Lambe (1905), Huene (1909) and Lambe (1911); to Predentata by von Zittel (1911) and Gilmore (1919); to Thyreophora by Nopcsa (1923) and Nopcsa (1923); to Thyreophoroidea by Huene (1934); to Ornithischia by Lapparent (1947); to Ponderopoda by Huene (1948) and Huene (1959); to Ceratopsia by Marsh (1890), Marsh (1895), Marsh (1896), Hatcher et al. (1907), Matthew (1915), Gregory and Mook (1925), Russell (1930), Huene (1950), Kuhn (1964), Estes (1964), Romer (1966), Swinton (1970), Thulborn (1971), Ostrom (1980), McIntosh (1981), Sues and Galton (1982), Russell (1984), Maryanska and Osmólska (1985), Bakker (1986), Carroll (1988) and Nessov (1995); to Ceratopsomorpha by Wolfe and Kirkland (1998); to Neoceratopsia by Dodson and Currie (1990), Dodson (1990), Chinnery and Weishampel (1998), Penkalski and Dodson (1999), Peng et al. (2001), Chinnery (2004), Ryan and Evans (2005) and Ryan (2007); and to Ceratopsoidea by Hay (1902), Zhao (1983), Norman (1984), Cooper (1985), Sereno (1986), Sereno (1997), Sereno (1998), Sereno (1999), Sereno (2000), Makovicky (2001), You and Dodson (2003) and Chinnery and Horner (2007).

Subtaxa

Centrosaurinae, Ceratopsinae, Ceratopsipes, Chasmosaurinae (syn. Chasmosaurus kaiseni), Monocloniinae, Pachyrhinosaurinae, Proceratops, Turanoceratops

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Synonyms

Ceratops montanus was named by Marsh (1888). Its type specimen is USNM 2411, a skull, and it is not a trace fossil. Its type locality is Ceratops type site, Cow Creek, which is in a Campanian terrestrial horizon in the Judith River Formation of Montana. It is the type species of Ceratops, Proceratops.

It was recombined as Proceratops montanus by Lull (1906); it was considered a nomen dubium by Dodson and Currie (1990), Dodson (1990) and Penkalski and Dodson (1999); it was considered a nomen dubium by Dodson et al. (2004), Ryan (2007) and Ryan et al. (2010).
Ceratops was named by Marsh (1888). It is not extant. Its type is Ceratops montanus. It is the type genus of Ceratopsidae, Ceratopsoidea.

It was synonymized subjectively with Monoclonius by Baur (1891); it was synonymized subjectively with Agathaumas by Lydekker (1893); it was replaced with Proceratops by Lull (1906); it was considered a nomen dubium by Dodson and Currie (1990), Dodson (1990), Lehman (1990), Dodson et al. (2004) and Ryan (2007).

It was assigned to Ceratopidae by Mansel-Pleydell (1902); to Ceratopsinae by Huene (1950); and to Ceratopsidae by Marsh (1888), Marsh (1888), Marsh (1895), Depéret (1896), Marsh (1896), Hay (1902), Hatcher et al. (1907), Huene (1909), Lambe (1911), von Zittel (1911), Matthew (1915), Gilmore (1919), Gregory and Mook (1925), Lull (1933), Romer (1956), Kuhn (1964), Romer (1966) and Swinton (1970).
Polyonax was named by Cope (1874).

It was synonymized subjectively with Trachodon by Hatcher (1902); it was synonymized subjectively with Agathaumas by Hay (1902) and Romer (1956); it was synonymized subjectively with Triceratops by Kuhn (1964) and Romer (1966); it was considered a nomen dubium by Hatcher et al. (1907); it was considered a nomen dubium by Dodson and Currie (1990) and Dodson et al. (2004).

It was assigned to Agathaumidae by Cope (1889), Cope (1889) and Cope (1891); to Ceratopsidae by Marsh (1895) and Marsh (1896); and to Reptilia by Huene (1909).
Agathaumas was named by Cope (1872). It is not extant. Its type is Agathaumas sylvestris.

It was synonymized subjectively with Triceratops by Romer (1966); it was considered a nomen dubium by Dodson and Currie (1990) and Dodson et al. (2004).

It was assigned to Dinosauria by Cope (1872); to Madrosauridae by Sauvage (1882); to Hadrosauridae by Dollo (1882), Marsh (1882), Marsh (1884) and Marsh (1885); to Scelidosauria by Seeley (1892); to Agathaumidae by Cope (1889), Cope (1891), Cope (1892), Cope (1898) and Lull (1906); and to Ceratopsidae by Marsh (1895), Marsh (1896), Hay (1902), Hatcher et al. (1907), Huene (1909), von Zittel (1911), Russell (1930), Romer (1956) and Kuhn (1964).
Agathaumas sylvestris was named by Cope (1872). Its type specimen is AMNH 4000, a skull, and it is not a trace fossil. Its type locality is Black Buttes Station, which is in a Maastrichtian estuary/bay sandstone/coal in the Lance Formation of Wyoming. It is the type species of Agathaumas.

It was considered a nomen dubium by Dodson and Currie (1990) and Dodson et al. (2004).
Bison alticornis was named by Marsh (1887). It is not extant. Its type specimen is USNM 4739. Its type locality is Windsor Addition, Highland, which is in a Maastrichtian terrestrial sandstone in the Denver Formation of Colorado.

It was recombined as Polyonax alticornis by Cope (1889); it was recombined as Ceratops alticornis by Marsh (1889) and Hay (1902); it was recombined as Triceratops alticornis by Hatcher et al. (1907) and Kuhn (1964); it was considered a nomen dubium by Lull (1933), Dodson and Currie (1990), Ostrom and Wellnhofer (1990) and Dodson et al. (2004).
Polyonax mortuarius was named by Cope (1874). Its type specimen is USNM 244535/AMNH 3950 and is not a trace fossil. Its type locality is Bijou Creek (USG&GS), which is in a Maastrichtian terrestrial horizon in the Laramie Formation of Colorado.

It was recombined as Agathaumas mortuarius by Hay (1902); it was recombined as Triceratops mortuarius by Kuhn (1964); it was considered a nomen dubium by Hatcher et al. (1907), Dodson and Currie (1990) and Dodson et al. (2004).
Agathaumidae was named by Cope (1889).

It was synonymized subjectively with Ceratopsidae by Lambe (1902), Hay (1902) and Hatcher et al. (1907).

It was assigned to Dinosauria by Cope (1889); to Iguanodontia by Baur (1891); and to Orthopoda by Cope (1889), Cope (1891) and Cope (1898).
Triceratops maximus was named by Brown (1933). Its type specimen is AMNH 5040, a set of vertebrae, and it is not a trace fossil. Its type locality is spring on Rock Creek (AMNH), which is in a Maastrichtian terrestrial sandstone in the Hell Creek Formation of Montana.

It was considered a nomen dubium by Lull (1933); it was considered a nomen dubium by Dodson and Currie (1990), Ostrom and Wellnhofer (1990) and Dodson et al. (2004).
Triceratops galeus was named by Marsh (1889). Its type specimen is USNM 2410. Its type locality is Brighton area, Henderson (1), which is in a Maastrichtian terrestrial horizon in the Arapahoe Formation of Colorado.

It was considered a nomen dubium by Lull (1933); it was considered a nomen dubium by Dodson and Currie (1990), Ostrom and Wellnhofer (1990) and Dodson et al. (2004).
Ugrosaurus was named by CoBabe and Fastovsky (1987). It is not extant. Its type is Ugrosaurus olsoni.

It was considered a nomen dubium by Dodson and Currie (1990), Lehman (1990) and Dodson et al. (2004).

It was assigned to Ceratopsidae by CoBabe and Fastovsky (1987).
Ugrosaurus olsoni was named by CoBabe and Fastovsky (1987). Its type specimen is UCMP 128561, a skull, and it is not a trace fossil. Its type locality is Dave's Nose, UCMP V-83224, which is in a Maastrichtian fluvial-lacustrine siltstone in the Hell Creek Formation of Montana. It is the type species of Ugrosaurus.

It was considered a nomen dubium by Dodson and Currie (1990) and Dodson et al. (2004).
Triceratops ingens was named by Lull (1915). Its type specimen is YPM 1828. Its type locality is south of Dogie Creek (YPM 1828), which is in a Maastrichtian terrestrial horizon in the Lance Formation of Wyoming.

It was considered a nomen dubium by Dodson and Currie (1990); it was considered a nomen nudum by Ostrom and Wellnhofer (1990); it was considered a nomen dubium by Dodson et al. (2004).
Pachyrhinosauridae was named by Sternberg (1950).

It was synonymized subjectively with Ceratopsidae by Langston (1967).

It was assigned to Ceratopsia by Sternberg (1950), Romer (1956), Kuhn (1964) and Romer (1966).

Synonymy list

Year	Name and author
1872	Agathaumas Cope pp. 481-483
1872	Agathaumas sylvestris Cope pp. 481-483
1873	Agathaumas sylvestris Cope p. 230
1874	Polyonax Cope
1874	Polyonax mortuarius Cope
1875	Agathaumas sylvestris Cope p. 435
1882	Agathaumas Dollo
1882	Agathaumas Marsh
1882	Agathaumas Sauvage p. 35
1884	Agathaumas Marsh p. 69
1885	Agathaumas Marsh p. 764
1887	Bison alticornis Marsh
1888	Ceratops Marsh
1888	Ceratops montanus Marsh p. 477
1888	Ceratops Marsh p. 478
1888	Ceratopsidae Marsh p. 478
1889	Agathaumas Cope
1889	Agathaumas sylvestris Cope
1889	Agathaumidae Cope
1889	Polyonax Cope
1889	Polyonax mortuarius Cope
1889	Agathaumidae Cope p. 16
1889	Polyonax alticornis Cope p. 290
1889	Agathaumidae Cope p. 715
1889	Polyonax Cope p. 906
1889	Ceratops alticornis Marsh pp. 174-175
1889	Triceratops galeus Marsh pp. 174-175
1890	Ceratopsidae Marsh p. 425
1891	Agathaumidae Baur p. 450
1891	Agathaumas Cope p. 43
1891	Agathaumas sylvestris Cope p. 43
1891	Agathaumidae Cope p. 43
1891	Polyonax Cope p. 43
1892	Agathaumas Cope p. 758
1892	Agathaumas sylvestris Cope p. 758
1892	Agathaumas Seeley
1895	Agathaumas Marsh
1895	Ceratops Marsh
1895	Ceratopsidae Marsh
1895	Polyonax Marsh
1896	Ceratopsidae Depéret p. 184
1896	Ceratops Depéret p. 185
1896	Agathaumas Marsh p. 243
1896	Ceratops Marsh p. 243
1896	Ceratopsidae Marsh p. 243
1896	Polyonax Marsh p. 243
1898	Agathaumas Cope p. 70
1898	Agathaumas sylvestris Cope p. 70
1898	Agathaumidae Cope p. 70
1902	Agathaumas Hay p. 498
1902	Agathaumas mortuarius Hay p. 498
1902	Agathaumas sylvestris Hay p. 498
1902	Ceratopsidae Hay p. 498
1902	Triceratops galeus Hay p. 498
1902	Ceratops Hay p. 500
1902	Ceratops alticornis Hay p. 500
1902	Ceratops montanus Hay p. 500
1902	Polyonax mortuarius Lambe p. 68
1902	Ceratopidae Mansel-Pleydell p. lxxv
1902	Ceratops Mansel-Pleydell p. lxxxiv
1902	Ceratops montanus Mansel-Pleydell p. lxxxv
1905	Ceratopsidae Lambe p. 49
1906	Agathaumas Lull p. 144
1906	Proceratops montanus Lull p. 144
1907	Ceratopsidae Hatcher et al. p. 10
1907	Agathaumas Hatcher et al. p. 11
1907	Ceratops Hatcher et al. p. 11
1907	Agathaumas sylvestris Hatcher et al. p. 168
1907	Triceratops alticornis Hatcher et al. p. 170
1907	Ceratops montanus Hatcher et al. p. 172
1909	Ceratopsidae Huene p. 17
1909	Agathaumas Huene p. 18
1909	Ceratops Huene p. 18
1909	Polyonax Huene p. 18
1911	Ceratops Lambe p. 9
1911	Ceratopsidae Lambe p. 9
1911	Ceratopsidae von Zittel p. 295
1911	Agathaumas von Zittel p. 296
1911	Ceratops von Zittel p. 296
1915	Ceratops montanus Lambe p. 10
1915	Triceratops ingens Lull
1915	Ceratops Matthew p. 30
1915	Ceratopsidae Matthew p. 32
1919	Ceratops Gilmore p. 64
1919	Ceratopsidae Gilmore p. 64
1923	Ceratopsidae Nopcsa p. 106
1925	Ceratopsidae Gregory and Mook p. 4
1925	Ceratops Gregory and Mook p. 5
1930	Agathaumas Russell p. 140
1930	Agathaumas sylvestris Russell p. 140
1930	Ceratopsidae Russell p. 140
1930	Ceratops montanus Russell p. 142
1933	Triceratops maximus Brown
1933	Ceratops Lull p. 96
1933	Ceratops montanus Lull p. 96
1934	Ceratopsidae Huene p. 184
1947	Ceratopsidae Lapparent pp. 17-18
1948	Ceratopsidae Huene p. 95
1950	Ceratops Huene p. 351
1950	Ceratopsidae Huene p. 351
1950	Pachyrhinosauridae Sternberg p. 109
1956	Agathaumas Romer p. 638
1956	Ceratops Romer p. 639
1956	Pachyrhinosauridae Romer p. 639
1959	Ceratopsidae Huene p. 122
1964	Ceratopsidae Estes p. 145
1964	Agathaumas Kuhn p. 54
1964	Agathaumas silvestris Kuhn p. 54
1964	Ceratopsidae Kuhn p. 54
1964	Ceratops Kuhn p. 55
1964	Ceratops montanus Kuhn p. 55
1964	Triceratops mortuarius Kuhn p. 59
1964	Pachyrhinosauridae Kuhn p. 60
1964	Triceratops alticornis Kuhn p. 60
1964	Triceratops galeus Kuhn p. 60
1964	Triceratops ingens Kuhn p. 60
1964	Triceratops maximus Kuhn p. 60
1964	Ceratops montanus Russell p. 10
1964	Triceratops maximus Russell p. 17
1966	Ceratops Romer p. 371
1966	Ceratopsidae Romer p. 371
1966	Pachyrhinosauridae Romer p. 371
1970	Ceratops Swinton p. 254
1970	Ceratopsidae Swinton p. 254
1971	Ceratopsidae Thulborn pp. 77-78 fig. 4
1980	Ceratopsidae Ostrom p. 23
1981	Ceratopsidae McIntosh p. 41
1982	Ceratopsidae Sues and Galton p. 189
1983	Ceratopsidae Zhao p. 300
1984	Ceratopsidae Norman p. 160 fig. 2
1984	Ceratopsidae Russell p. 29
1985	Ceratopsidae Cooper pp. 292-293
1985	Ceratopsidae Maryanska and Osmólska pp. 143-144 fig. 1
1986	Ceratopsidae Bakker p. 461
1986	Ceratopsidae Sereno
1987	Ugrosaurus CoBabe and Fastovsky p. 149 figs. 1-4
1987	Ugrosaurus olsoni CoBabe and Fastovsky p. 149
1988	Ceratopsidae Carroll
1990	Ceratopsidae Dodson p. 578
1990	Ceratopsidae Dodson and Currie p. 611
1995	Ceratopsidae Nessov p. 54
1997	Ceratopsidae Sereno p. 443
1998	Ceratopsidae Chinnery and Weishampel pp. 578-579 fig. 10
1998	Ceratopsidae Sereno p. 62
1998	Ceratopsidae Wolfe and Kirkland p. 307 fig. 5
1999	Ceratopsidae Penkalski and Dodson
1999	Ceratopsidae Sereno p. 2138 fig. 2
2000	Ceratopsidae Sereno p. 482
2001	Ceratops montanus Horner et al. p. 10
2001	Ceratopsidae Makovicky p. 254 fig. 18.3
2001	Ceratopsidae Peng et al. p. 35
2003	Ceratopsidae You and Dodson
2004	Ceratopsidae Chinnery pp. 584-585 fig. 9
2005	Ceratopsidae Ryan and Evans p. 323
2007	Ceratopsidae Chinnery and Horner p. 638 fig. 6
2007	Ceratopsidae Ryan p. 391 fig. 12

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