Mammalia - Perissodactyla - Brontotheriidae

Full reference: H. F. Osborn. 1908. New or little known titanotheres from the Eocene and Oligocene. Bulletin of the American Museum of Natural History 24(32):599-617

Parent taxon: Dolichorhininae according to B. J. Mader 1998

Sister taxa: Metarhinus, Sphenocoelus

Subtaxa: Mesatirhinus megarhinus

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Ecology: ground dwelling browser