Insecta - Coleoptera - Geotrupidae

Parent taxon: Geotrupidae according to J. F. Zhang 1989

Sister taxa: Bolboceratinae, Coprologus, Geotrupinae, Lethrinae, Orrhodomala, Typhaeus

Subtaxa: Geotrupes atavus, Geotrupes jiaoyanshanense, Geotrupes messelensis

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Ecology: