Coudoulous I (Faunal Units IV and V)
Basic information
Sample name: Coudoulous I (Faunal Units IV and V)

Reference: P. Fernandez, M. Jeannet, J. Jaubert, and J.-P. Brugal. 2021. Paleoecological first results on the Middle Pleistocene sequence of Coudoulous I (Quercy, Lot, France). Alpine and Mediterranean Quaternary 34(1):89-107 [ER 3857]
Geography
Country: France


Coordinate: 44° 28' 20" N, 1° 39' 50" E
Coordinate basis: stated in text

Time interval: Middle Pleistocene

Max Ma: 0.229

Min Ma: 0.1613

Age basis: other

Geography comments: "in the Quercy area... at the junction of the river LOt and its tributary, the Célé"
the entire faunal sequence is bracketed by an ESR/U-series date of 161.3 +/- 14 kyr at the top and two U/Th dates of 229 + 78 - 42 and 202 + 78 - 39 kyr at the base

Environment
Lithology: claystone

Taphonomic context: cave,human accumulation,pitfall trap

Archaeology: stone tools

Habitat comments: a pitfall trap with fossils from "clays, sands, angular limestone rocks"
there is a "very rich archaeological level" immediately above the fossiliferous sequence that dates to the "Early Middle Paleolithic" and there are "scarce and dispersed artefacts of the same age" in the fossiliferous levels
the site "was used for marginal scavenging by Lower Paleolithic hominid groups and as a kill-butchery site during the EMP", so it is at least partially a human accumulation
from stratigraphic units 7c, 7d, 7x, 7e, and 7f
there is apparently no sandstone in this interval, so the fossils are presumably from clays

Methods
Life forms: bats,carnivores,rodents,ungulates,other small mammals,lizards,snakes,frogs,salamanders

Sampling methods: quarry,screenwash

Sample size: 3226 specimens

Years: 1994 - 2003

Sampling comments: there was a "rescue excavation" in 1978 - 1980, but the main excavation was between 1994 and 2003
the high abundance of micromammals is taken to mean that screening was used

Metadata
Sample number: 4225

Contributor: John Alroy

Enterer: John Alroy

Created: 2023-04-14 17:52:54

Modified: 2023-04-14 07:52:54

Abundance distribution
96 species
18 singletons
total count 3226
geometric series index: 148.8
Fisher's α: 18.599
geometric series k: 0.9357
Hurlbert's PIE: 0.9221
Shannon's H: 3.0879
Good's u: 0.9944
Each square represents a species. Square sizes are proportional to counts.
Register
Microtus arvalis553
Microtus agrestis74
Stenocranius gregalis265
"Microtus gregalis"
Alexandromys oeconomus4336.5 g
"Microtus oeconomus"
Chionomys nivalis3
Microtus brecciensis68
Microtus multiplex26
Microtus gerbii9
"Microtus pyrenaicus"
Microtus subterraneus30
Myodes glareolus28
"Clethrionomys glareolus"
Lemmus lemmus1
Dicrostonyx torquatus252.0 g
Pliomys lenki22
Pliomys chalinei1
Arvicola amphibius2
"Arvicola terrestris"
Arvicola cantiana15
Allocricetus bursae165
Apodemus sylvaticus293
Apodemus flavicollis7
Apodemus uralensis21
Eliomys quercinus400
Muscardinus avellanarius4
Sciurus vulgaris1311 g
Erinaceus europaeus3
Erinaceus davidi4
Talpa europaea9
Talpa minor1
Sorex araneus6
Sorex minutus7
Sorex minutissimus1
Crocidura russula6
Crocidura leucodon1
Crocidura suaveolens1
Castor cf. fiber1
Oryctolagus cuniculus62.5 kg browser-grazer
Lepus europaeus83.4 kg herbivore
Lepus timidus5
Ochotona pusilla7
Rhinolophus ferrumequinum120.0 g insectivore
Rhinolophus hipposideros15.3 g
Myotis myotis325.7 g
Myotis blythii224.6 g
Myotis capaccinii110.0 g
Myotis mystacinus25.5 g
Myotis bechsteinii18.7 g
Vespertilio murinus111.9 g
Eptesicus serotinus122.6 g
Nyctalus noctula129.2 g insectivore
Pipistrellus pipistrellus24.4 g insectivore
Barbastella barbastellus38.0 g insectivore
Plecotus auritus57.6 g
Plecotus austriacus38.3 g insectivore
Salamandra salamandra4
Pelobates fuscus2
Bufo bufo6
Bufo calamita6
Hyla arborea2
Rana dalmatina4
Rana temporaria3
Lacerta agilis2
Lacerta viridis8
Podarcis cf. muralis3
Chalcides chalcides2
Anguis fragilis8
Coluber viridiflavus3
Coronella austriaca6
Coronella girondica6
Elaphe longissima2
Rhinechis scalaris1
Malpolon monspessulanus2
Natrix maura2
Natrix natrix6
Vipera aspis3
Vipera berus5
Vipera ursinii3
Cervus elaphus114104 kg
Dama dama1462 kg
Capreolus capreolus1522 kg browser
Bison priscus99
compared to subspecies mediator
Equus cf. mosbachensis39
Hemitragus cedrensis50
Palaeoloxodon antiquus23
Stephanorhinus cf. hemitoechus8
Sus scrofa154 kg herbivore
Canis lupus lupus28943 kg carnivore
Cuon alpinus1614 kg carnivore
Vulpes vulpes95.3 kg carnivore-insectivore
Felis silvestris24.5 kg carnivore
Lynx lynx216 kg carnivore
Panthera spelaea96
Ursus arctos117156 kg frugivore-carnivore
Panthera pardus233 kg carnivore
Mustela erminea1121 g
Mustela nivalis10277.5 g carnivore
Mustela putorius4681 g carnivore
Meles meles811 kg invertivore-herbivore