Coudoulous I (Faunal Unit VI)
Basic information
Sample name: Coudoulous I (Faunal Unit VI)

Reference: P. Fernandez, M. Jeannet, J. Jaubert, and J.-P. Brugal. 2021. Paleoecological first results on the Middle Pleistocene sequence of Coudoulous I (Quercy, Lot, France). Alpine and Mediterranean Quaternary 34(1):89-107 [ER 3857]
Geography
Country: France

Coordinate: 44° 28' 20" N, 2° 40' 50" E
Latlng basis: stated in text

Time interval: Middle Pleistocene

Max Ma: 0.229

Min Ma: 0.1613

Age basis: other

Geography comments: "in the Quercy area... at the junction of the river LOt and its tributary, the Célé"
the entire faunal sequence is bracketed by an ESR/U-series date of 161.3 +/- 14 kyr at the top and two U/Th dates of 229 + 78 - 42 and 202 + 78 - 39 kyr at the base

Environment
Lithology: sandstone

Taphonomic context: cave, human accumulation, pitfall trap

Archaeology: stone tools

Habitat comments: a pitfall trap with fossils from "clays, sands, angular limestone rocks"
there is a "very rich archaeological level" immediately above the fossiliferous sequence that dates to the "Early Middle Paleolithic" and there are "scarce and dispersed artefacts of the same age" in the fossiliferous levels
the site "was used for marginal scavenging by Lower Paleolithic hominid groups and as a kill-butchery site during the EMP", so it is at least partially a human accumulation
from stratigraphic units 8a, 8b, 8b'1, and 8b'2, the first two based on Fig. 2 being a sandstone that is presumably the source of much or perhaps most of the material because it does not include breccia

Methods
Life forms: bats, carnivores, rodents, ungulates, other small mammals, lizards, snakes, frogs, salamanders

Sampling methods: quarry, screenwash

Sample size: 823 specimens

Years: 1994 - 2003

Net or trap nights: 0

Basal area status: not applicable

Sampling comments: there was a "rescue excavation" in 1978 - 1980, but the main excavation was between 1994 and 2003
the high abundance of micromammals is taken to mean that screening was used

Metadata
Sample no: 4226

Contributor no: John Alroy

Enterer: John Alroy

Created: 2023-04-14 21:05:18

Modified: 2023-04-14 11:05:18

Abundance distribution
Each square represents a species. Square sizes are proportional to counts. Values are logged.
Statistics
57 species
14 singletons
total count 823
geometric series index: 99.4
Fisher's α: 13.913
geometric series k: 0.904
Hurlbert's PIE: 0.852
Shannon's H: 2.7404
Good's u: 0.983
Register
Microtus arvalis 285
Microtus agrestis 81
Stenocranius gregalis 10
"Microtus gregalis"
Alexandromys oeconomus 2
"Microtus oeconomus"
Chionomys nivalis 1
Microtus brecciensis 28
Microtus multiplex 22
Microtus gerbii 13
"Microtus pyrenaicus"
Microtus subterraneus 19
Clethrionomys glareolus (bank vole)28
Pliomys lenki 11
Arvicola cantiana 3
Allocricetus bursae 4
Apodemus sylvaticus 27
Apodemus uralensis 12
Eliomys quercinus 27
Muscardinus avellanarius 2
Talpa europaea (European mole)5
Sorex araneus 5
Sorex minutus 5
Crocidura russula (greater white-toothed shrew)5
Lepus europaeus (European hare)23.4 kg
Ochotona pusilla (steppe pika)1
Myotis myotis 225.7 g
Myotis blythii 124.6 g
Pipistrellus pipistrellus 14.4 g
Plecotus auritus 27.6 g
Plecotus austriacus 18.3 g
Triturus helveticus 2
Bufo bufo 1
Lacerta agilis 4
Lacerta viridis 2
Podarcis cf. muralis 3
Anguis fragilis 6
Coluber viridiflavus 2
Coronella austriaca 1
Coronella girondica 1
Elaphe longissima 1
Malpolon monspessulanus 1
Natrix maura 1
Natrix natrix 2
Vipera aspis 1
Vipera berus 2
Vipera ursinii 2
Cervus elaphus (red deer)7
Capreolus capreolus (roe deer)1
Bison priscus (bison)68
compared to subspecies mediator
Equus cf. mosbachensis (horse)7
Hemitragus cedrensis (tahr)14
Sus scrofa (pig)2
Canis lupus lupus (gray wolf)1843.0 kg
Vulpes vulpes (red fox)1
Lynx lynx (Eurasian lynx)6
Panthera spelaea (cave lion)2
Ursus arctos (brown bear)48
Mustela nivalis (least weasel)961.7 g
Meles meles (European badger)311.0 kg