Holly's Microsite
Basic information
Sample name: Holly's Microsite

Sample aka: SC-188

Reference: K. D. Rose. 1981. The Clarkforkian land-mammal age and mammalian faunal composition across the Paleocene-Eocene boundary. University of Michigan Papers on Paleontology 26:1-197 [ER 4102]
Geography
Country: United States

State: Wyoming


County: Park


Coordinate: 44.85° N, -109.10° W
Coordinate basis: stated in text

Formation: Willwood

Time interval: Late Palaeocene

Zone: Clarkforkian

Section: Polecat Bench

Unit number: 7

Max Ma: 57.101

Min Ma: 56

Age basis: other

Geography comments: "SE 1/4, S 1, T 56 N, R 102 W, and SW 1/4, S 6, T 56 N, R 101 W (Sec. 4)"
assigned to the Plesiadapis cookei zone of the Clarkforkian, meaning the Clarkforkian part of the lower Willwood Formation, which is entirely in chron 24r (see for example Gingerich 2001)
the meter level is said to be 1280 m by Gunnell and Gingerich (1991) and appears to be 1260 m in Fig. 3 of Krause (1986)
the age assignments are the base of chron 24r according to Ogg (2020) and the Palaeocene-Eocene boundary according to the ICS 2003 time scale

Environment
Lithology: mudstone

Habitat comments: "Gray mudstone at the base of this locality contains a pocket that, through quarrying and screen-washing, has yielded the richest Clarkforkian collection from a single site in the Clark's Fork Basin" but several "larger specimens" have "been excavated from drab mudstones slightly above the level of the quarry"

Methods
Life forms: carnivores,primates,rodents,ungulates,other large mammals,other small mammals

Sampling methods: quarry,screenwash

Sample size: 472 specimens

Years: 1977

Museum: UMMP

Sampling comments: "discovered in 1977 by B. Holly Smith"
"we found surface concentrations of small vertebrates, screenwashing was performed" and was "highly successful" and the site was quarried "extensively": "About 30 sacks (approximately one ton) of matrix have been washed"
Secord (2008) reports a much larger sample from this site with different counts and more species that I have integrated into Rose's version
"Phenacodus, Coryphodon, and Dipsalidictis (Oxyaena) aequidens appeared in a faunal list for Holly’s Microsite published by Krause (1986) but are excluded here because they appear to have been surface collected from the surrounding area (SC-188), based on museum records"
the omitted Phenacodus primaevus of Rose is the same as Phenacodus intermedius of Thewissen (2008)
one specimen of Dissacus sp. is also omitted by Secord (2008), and I am unable to match this identification to his list
see also Krause (1986)

Metadata
Sample number: 4531

Contributor: John Alroy

Enterer: John Alroy

Modifier no: John Alroy

Created: 2024-12-01 04:27:32

Modified: 2024-12-06 22:52:31

Abundance distribution
37 species
8 singletons
total count 472
geometric series index: 58.2
Fisher's α: 9.401
geometric series k: 0.8820
Hurlbert's PIE: 0.9044
Shannon's H: 2.8251
Good's u: 0.9831
Each square represents a species. Square sizes are proportional to counts.
Register
Microcosmodon rosei16
type
Ectypodus powelli92
Peradectes protinnominatus21
Krishtalka and Stucky (1983): "Peradectes cf. chesteri"
Prodiacodon cf. tauricinerei3
Palaeoryctes punctatus2
Secord (2008)
Labidolemur kayi8
Krause (1986): :"Apatemys sp."
Leptacodon rosei27
Gingerich (1987): "Leptacodon cf. packi"
Plagioctenodon cf. krausae8
Leipsanolestes cf. siegfriedti5
Secord (2008): "Leipsanolestes siegfriedti"
Diacodon cf. minutus = †Adunator cf. minutus18
Worlandia inusitata87
type
Niptomomys doreenae3
Tinimomys graybulliensis2
Plesiadapis cookei17
Niptomomys doreenae2
Tinimomys graybulliensis3
Carpolestes simpsoni31
Bloch and Gingerich (1998): "Carpolestes nigridens"
Phenacolemur cf. praecox17
Secord (2008): "Phenacolemur pagei"
Phenacolemur cf. simonsi1
Secord (2008)
Ignacius graybullianus7
Chriacus sp.2
Secord (2008)
Princetonia sp.4
Secord (2008)
Thryptacodon sp.1
Secord (2008): "Thryptacodon cf. antiquus"
Ectocion osbornianus9
Haplomylus simpsoni1
Esthonyx ancylion1
Arctostylops steini1
Didymictis proteus3
Polly (1997): "Didymictis protenus protenus"
Viverravus cf. bowni = †Viverravus cf. laytoni8
Krause (1986): "Viverravus sp." said to be new
Uintacyon rudis1
Tytthaena sp.1
Secord (2008): cf.
Paramys atavus = †Acritoparamys atavus32
apparently also includes 17 specimens of "Paramys sp."
Acritoparamys atwateri8
Secord (2008)
Microparamys cheradius19
Ivy (1990): one specimen of Rose''s is definitely this species: the type and others appear to have been collected later
Microparamys cf. minutus1
Secord (2008)
Paramys taurus7
(Secord (2008)
Reithroparamys sp.3
Ivy (1990)