Abundance: 0.09 to 48.77% (median 4.49%)

Latitudinal range: 29.2° to 43.1°

Habitats: (27), Mediterranean woodland (3), temperate broadleaf/mixed forest (3)

Found in 33 samples

Greece: Lake Prespa National Park, Milia (MIL5)

Iran: Ilam, Wezmeh Cave, Tappeh Sang-e Chakhmaq (West Mound), Tappeh Sang-e Chakhmaq (East Mound), Qasr-e Ahmad

Israel: Qesem Cave (Concentration 1), Qesem Cave (Concentration 2), Manot Cave (Area D, Biogenic Unit), Motza (Early PPNB), Motza (Middle PPNB), Tel Ro'im West (Late PPNB), Tel Ro'im West (PPNC), Tel Ro'im West (PN), Yiftah'el, Marj Rabba, El-Wad Cave (Early Natufian), El-Wad Cave (Late Natufian), Qesem Cave Hearth unit, Qesem Cave LSBS, Ein Gev I

Italy: Poggi di Prata, Capalbiaccio (Building J, Level III)

Jordan: Dibbeen Nature Reserve

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E)

Montenegro: Great Ulcinj Beach

Morocco: Ifri El Baroud (Unit B2)

Portugal: Gruta de Figueira Brava (Area C)

Spain: Bolomor Cave (Level XI), Bolomor Cave (Level IV)

Turkey: Madran Mountain, Direkli Cave

See also Testudo brevitesta, Testudo graeca, Testudo hermanni, Testudo sp.