Abundance: 0.30 to 18.93% (median 2.50%)

Latitudinal range: 38.5° to 43.1°

Habitats: (3), Mediterranean woodland (1), temperate broadleaf/mixed forest (2)

Found in 6 samples

Greece: Lake Prespa National Park

Italy: Poggi di Prata

Montenegro: Great Ulcinj Beach

Portugal: Gruta de Figueira Brava (Area C)

Spain: Bolomor Cave (Level XI), Bolomor Cave (Level IV)

See also Testudo, Testudo brevitesta, Testudo graeca, Testudo sp.