Napu Valley (natural forest) Thirunelly Forests Napu Valley (young secondary forest) Napu Valley (agroforestry ecosystem) Napu Valley (annual culture) Ba Be National Park (dung beetles) Danum Valley Field Centre (Riverine A) Sungkai Reserve Forest (secondary forest) The Ecological Register
global
North America
United States
South America/Africa
Europe
Eurasia
Africa/Australasia
•
•
•
•
•
•
•
•
Onthophagus pacificus (scarab beetle): 8 samples
Standardised estimates are produced by subsampling and then extrapolating.
Raw richness may be higher when an abundance distribution is uneven.