Blackbeard Island Cumberland Island East of Arapaho Prairie Arapaho Prairie Welder Wildlife Refuge (open chaparral) Welder Wildlife Refuge (1985) North of Caldwell (open pasture, 1979) Hutcheson Memorial Forest disturbed field (June/July) Hutcheson Memorial Forest old growth forest (June/July) Lake City Research Center Site A Lake City Research Center Site B Brookings County (cattle manure) Brookings County (sheep manure) Missouri Ozark Forest Ecosystem Project (no harvest management) The Ecological Register
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Onthophagus pennsylvanicus (scarab beetle): 14 samples
Standardised estimates are produced by subsampling and then extrapolating.
Raw richness may be higher when an abundance distribution is uneven.
Current reference: Shotwell 1968 (ER 3992)