Abundance: 0.00 to 95.31% (median 4.86%)

Latitudinal range: -40.6° to 67.8°

Habitats: (345), Mediterranean woodland (2), desert/xeric shrubland (6), temperate broadleaf/mixed forest (5), temperate coniferous forest (1), temperate savanna (1), tropical/subtropical dry broadleaf forest (2), tropical/subtropical moist broadleaf forest (1), tropical/subtropical savanna (8)

Found in 371 samples

Algeria: Ain Boucherit (Lower), Ain Boucherit (Upper)

Argentina: Acambuco, Empalme Querandíes 1, Paso Otero (pardo), Paso Otero (verde)

Armenia: Godedzor Level II B

Australia: Goonoo State Conservation Area (on-track, pre-baiting)

Belize: Extinction Cave

Botswana: Moritsane

Brazil: Lagoa das Abelhas, Jirau Paleontological Site, Curimatãs Paleontological Site, Catimbau National Park, Lage Grande Paleontological Site, Poço Redondo

Bulgaria: Redaka II

Canada: Old Crow River Loc. 11A (square S30E120)

Chile: Pilauco (lower PB-7), GNL Quintero 1

China: Altun Mountains, Mount Kalamaili Ungulate Nature Reserve, Yanchiwan National Nature Reserve, Shuidonggou Locality 12, Dazhushan, Songhua River, Tawendaliha, Xiariyamakebu, Talitaliha, Gezishan Locality 10 (Fourth layer), Wanglang Nature Reserve (2020), Foping National Nature Reserve (2020), Yujiagou (Layer 3B), Yujiagou (Layer 4), Jinsitai Cave (Layer 7), Jinsitai Cave (Layer 8), Guandimiao, Shuangta, Muzhuzhuliang, Huoshiliang, Nanshan, Dabeigou, Shuidonggou Locality 7, Lower Pubu Cave, Xiaonanshan

Croatia: Mujina Pecina (Level B), Mujina Pecina (Level D1), Mujina Pecina (Level D2)

Democratic Republic of the Congo: Katanda 9

Ecuador: Tanque Loma, Refugio de Vida Silvestre Marina y Costera Pacoche, Rio Engabao

Egypt: Gebel Silsila 2B-I (Afian), Gebel Silsila 2B (Survey), Berenike Trogodytika (fort), Berenike Trogodytika (dump), Berenike Trogodytika (gate)

Eritrea: Uadi Aalad, Dandiero

Ethiopia: Bulbula (B1s1), Porc-Epic Cave, Harlaa, Bodo D'ar, Andalee, Simbiro III

France: Grotte du Bison (Level I), Mas d'Azil, Les Auzières 2, Les Canalettes (Layer 2), Les Canalettes (Layer 3), Les Canalettes (Layer 4), Noisetier Cave, Chez-Pinaud Jonzac (Layer 22), Coudoulous I (Faunal Unit II), Coudoulous I (Faunal Unit III), Coudoulous I (Faunal Units IV and V), Coudoulous I (Faunal Unit VI), Coudoulous I (Faunal Unit VII), Bois-de-Riquet (Locus 1), Caune de l'Arago (Unit D), Saint-Vallier (LD3), Vergisson II, Romain-la-Roche, Lazaret (C II supérieur UA 13-17), Payre (ensemble G), Payre (ensemble F), Payre (ensemble D), Orgnac 3 (level 1), Les Pradelles (level 9), Les Pradelles (level 10), Tourville-la-Rivière (level D1), Lunel-Viel (level 9), Soucy 1

Georgia: Dzudzuana Cave (lower area, Unit B)

Germany: Salzgitter Lebenstedt, Untermassfeld, Schöningen 13 II-4

Greece: Vatera (F-site), Kalamotó (KAL), Kalamotó (KLT)

Guatemala: Chivacabé

Honduras: La Majadita

Hungary: Oltovány Castle

India: Takalghat, Naikund, Malvan (Period I), Malvan (Period II), Kunal - Period IB (Early Harappan - Layer 5), Mitathal (Layer 2), Chandankhera (Period IV: Vakataka), Charnel House Cave (MIS 3), Cathedral Cave (MIS 5), Gopnath

Iran: Wezmeh Cave, Kobeh Cave, Tappeh Sang-e Chakhmaq (East Mound)

Iraq: Palegawra Cave

Israel: Manot Cave (Area D, Biogenic Unit), Qubur al-Walaydah (Field 2), Revadim Quarry, Ubeidiya (Layer II-23), Revadim Quarry Area B, Qesem Cave Hearth unit, Qesem Cave LSBS, Ein Gev I, Oumm Qatafa Cave (layer D2)

Italy: Western Trento Province (2012), Cava Muracci, San Teodoro Cave (Unit B-I), San Teodoro Cave (Unit B-II), Canale delle Acque Alte (Layer D-E3), Canale delle Acque Alte (Layer C2), Canale delle Acque Alte (Layer C1), La Sassa Cave, Contrada Castro (Period B), Casal Selce 2, Pirro Nord 13, Capalbiaccio (Building J, Level III), Grotta Cucgiliana, Pantalla, Casal de'Pazzi, Poggio Rosso

Jordan: Ayn Qasiyya (Area A), Shishan Marsh 1 (Level 7b), Khirbat al-Mudayna al-'Aliya

Kazakhstan: Tasbas (Phase 2b), Tasbas (Phase 3), Begash (Phase 1b), Begash (Phase 2), Begash (Phase 3a), Begash (Phase 3b), Begash (Phase 4)

Kenya: Karungu, Mfangano Island, Kibogo, Lainyamok

Kyrgyzstan: Chap I

Lesotho: Sehonghong (GAP), Sehonghong (GWA), Sehonghong (ALP), Sehonghong (SA), Sehonghong (BARF), Sehonghong (RF), Sehonghong (RBL/CLBRF), Sehonghong (BAS), Sehonghong (MOS)

Malawi: Malowa

Mauritania: Dhar Tichitt Site 38, Dhar Tichitt Site 46

Mexico: Barranca de San Mateo Huexoyucán (Localidad 6)

Mongolia: Tolbor-21 (AH4), Otson Tsokhio and Zuun Shovkh, Zaraa Uul (Locus 2), Zaraa Uul (Locus 3)

Montenegro: Trlica (Bed 10), Trlica (Bed 5)

Morocco: Ifri El Baroud (Unit B2), Gazelle Cave (lower level), Gazelle Cave (upper level), Zouhrah Cave (L s1), Zouhrah Cave (L 1), Zouhrah Cave (L 2), Zouhrah Cave (L 3), Grotte des Rhinocéros (lower set), Grotte des Rhinocéros (upper set), Thomas Quarry Hominid Cave

Namibia: Waterberg Plateau Park (surrounding flatlands), Namib-Naukluft National Park, Etosha Region

North Korea: Chongphadae Cave

Panama: El Hatillo

Poland: Borsuka Cave (Layer VI), Raj Cave (layer 4), Raj Cave (layer 6), Sadowie 2 (Neolithic), Sadowie 2 (Early Bronze Age), Zabia Cave (Layer 17)

Portugal: Foz do Enxarrique, Gruta de Figueira Brava (Area C)

Romania: Noviodunum (TPF sector, Getic period), Noviodunum (TPF sector, Roman period), Noviodunum (TPF sector, Middle Ages)

Russia: Svetly (layer 5), Geographical Society Cave, Zapovednaya, Gornovo, Bugul'deika II Layer II-2, Sagan-Zaba II (Layer 1), Sagan-Zaba II (Layer 2), Sagan-Zaba II (Layer 3A), Sagan-Zaba II (Layer 3B), Bol'shoj Naryn (Paleosol Layers), Chagyrskaya Cave (Layer 5), Chagyrskaya Cave (Layer 6), Kamenka, Cheremukhovo-1 (Bed 8), Cheremukhovo-1 (Bed 9), Cheremukhovo-1 (Bed 10), Cheremukhovo-1 (Bed 11), Imanay Cave (Pleistocene Layer 1), Imanay Cave (Pleistocene Layer 2), Imanay Cave (Pleistocene Layer 4), Khayrgas Cave - Layer 5 (1998–1999 pit), Khayrgas Cave - Layer 6 (1998–1999 pit), Byzovaya, Muhkai II (layer 80)

Saudi Arabia: Ti’s al Ghadah

Serbia: Garicin Grad (southeast corner tower, first phase), Garicin Grad (southeast corner tower, second phase)

South Africa: Groenefontein Nature Reserve (thicket vegetation), Kloof/Hillcrest, Sibudu Cave (Pinkish Grey Sand), Equus Cave (Unit 1A), Equus Cave (Unit 1B), Table Mountain National Park, Plovers Lake, Blombos Cave (Phase M3), Spitzkloof A Rockshelter (MIS 2), Diepkloof Rock Shelter (MSA - Post HP), Diepkloof Rock Shelter (MSA - Late HP), Diepkloof Rock Shelter (MSA - Intermediate HP), Diepkloof Rock Shelter (MSA - Jack), Diepkloof Rock Shelter (MSA - Still Bay), Diepkloof Rock Shelter (MSA - Pre-Still Bay Lynn), Diepkloof Rock Shelter (MSA - Mike), Diepkloof Rock Shelter (Lower MSA), Boomplaas Cave (Member BLD), Boomplaas Cave (Member BLA), Boomplaas Cave (Member BRL), Boomplaas Cave (Member CL), Boomplaas Cave (Member GWA), Boomplaas Cave (Member LP), Boomplaas Cave (Member LPC), Boomplaas Cave (Member YOL), Boomplaas Cave (Member BP), Boomplaas Cave (Member OLP), Boomplaas Cave (Member BOL), Boomplaas Cave (Member OCH), Bundu Farm, Equus Cave (Unit 2A), Equus Cave (Unit 2B), GD 1, Wonderwerk Cave (excavation 2, stratum 2), Haasgat Dumpsite, Cornelia-Uitzoek (1998 - 2010 collection), Cornelia-Uitzoek (old collection), Duinefontein 2 Horizon 2

Spain: El Rincon-1, Huéscar 1, Abric Romaní (Level Ja), Abric Romaní (Level Jb), Jou Puerta, El Horno Cave (Level 1), El Horno Cave (Level 2), Cueva del Angel, El Mirador Cave, Cueva de Nerja (Last Glacial Maximum), Cueva de Nerja (Tardiglacial), El Mirón Cave (Final Magdalenian), El Mirón Cave (Upper Magdalenian), El Mirón Cave (Middle Magdalenian), El Cierro Cave (Level C/D), Fortress of Vilars (sector 7/1), Fuente Nueva 3 (layer 2), Fuente Nueva 3 (layer 3), Fuente Nueva 3 (layer 5), Bolomor Cave (Level XI), Bolomor Cave (Level XVIIc), Bolomor Cave (Level XII), Bolomor Cave (Level IV), Venta Micena 4-I, Venta Micena 4-II, Barranc de la Boella (La Mina 2), Barranc de la Boella (El Forn 1), Barranc de la Boella (El Forn 2), Barranc de la Boella (El Forn 3), Barranc de la Boella (El Forn 4), Punta Lucero III (Level F), Ambrona (AS1), Barranco Léon, Gran Dolina (TD6.3), La Puebla de Valverde

Sudan: Gala Abu Ahmed, Khor Shambat 1 (Mesolithic), Khor Shambat 1 (Neolithic), HP766

Syria: Tell Tuneinir (Area 6 - Market)

Tanzania: Ruaha National Park (dry season, random placement), Mlele Forest Reserve, Tli’ika, Luxmanda, Kuumbi Cave (Trench 10, Phase 1b), Kuumbi Cave (Trench 10, Phase 2), Kuumbi Cave (Trench 10, Phase 3), Kuumbi Cave (Trench 10, Phase 4), Ewass Oldupa (Trench 2), Ewass Oldupa (Trench 3), Ewass Oldupa (Trench 5)

Tunisia: Doukanet el Khoutifa, Meninx

Turkey: Marmaris, Ciglikara Nature Reserve, Koycegiz Wildlife Development Reserve, Göbekli Tepe (Enclosure D), Sarikol Tepe, Faber Quarry

Ukraine: Prolom II, Mirnoe, Grot Skeliastyi (III-3B), Grot Skeliastyi (III-3), Grot Skeliastyi (III-3-x), Buran-Kaya III Layer 4, Maidanetske (2013 season)

United Arab Emirates: Early ed-Dur (50 BC - 150 AD)

United Kingdom: Lynford Quarry, Balderton Sand and Gravel

United States: North Fork of the Cache la Poudre River, Davis Mountain Preserve, Ardis, Zesch Cave, Crystal Ball Cave, Welsh Cave, Silver Creek, OMNH locality VI03, Vess Cave, Papago Springs Cave, Haystack Cave (lower levels), Saber-tooth Cave, Golf Course, Rancho La Brea (Pit 91), Harrodsburg Crevice, Ripplemead, Cueva Quebrada, First American Bank Site (Feature 1), Taylor Bayou (west bank), Jack Wade, Schuiling Cave, Sunshine Locality (Unit E), Schulze Cave (layer C1), Schulze Cave (layer C2), Bogue Chitto Creek, West of US 287, Kincaid Shelter, Cumberland Cave

Uruguay: La Paz bonebed

Uzbekistan: Kara-tepe, Kyzyltepa

Venezuela: El Breal de Orocual (ORS20)

Zambia: Leopard's Hill Cave (Pre-Nachikufan), Makwe Rockshelter 2

Zimbabwe: Ingwezi Game Management Project

See also Equus africanus, Equus algericus, Equus altidens, Equus apolloniensis, Equus capensis, Equus complicatus, Equus conversidens, Equus ferus, Equus grevyi, Equus hemionus, Equus hydruntinus, Equus kiang, Equus lambei, Equus leidyi, Equus livenzovensis, Equus major, Equus mauritanicus, Equus mexicanus, Equus mosbachensis, Equus neogeus, Equus niobrarensis, Equus numidicus, Equus occidentalis, Equus ovodovi, Equus quagga, Equus sanmeniensis, Equus scotti, Equus sivalensis, Equus sp., Equus sp. large, Equus sp. small, Equus spp., Equus steinheimensis, Equus stenonis, Equus suessenbornensis, Equus tabeti, Equus torralbae, Equus wuesti, Equus zebra