Abundance: 0.08 to 8.96% (median 0.85%)

Latitudinal range: -29.8° to 46.9°

Habitats: (15) only

Found in 15 samples

Botswana: Moritsane (Holocene)

Iraq: Jebel Hamrin Sud (Late Miocene)

Kenya: Kanapoi (Early Pliocene)

Lesotho: Sehonghong (GWA) (Holocene), Sehonghong (ALP) (Holocene)

Spain: Baza-1 (Early Pliocene), Caló den Rafelino (Early Pliocene)

Switzerland: Wallenried (Early Miocene)

Tanzania: Kuumbi Cave (Trench 10, Phase 1b) (Holocene)

Thailand: Spirit Cave (Layer 3) (Late Pleistocene)

Turkey: Ulucak Höyük (Level VI) (Holocene), Ulucak Höyük (Level V) (Holocene), Ulucak Höyük (Level IV) (Holocene)

United States: Farish Ranch (Middle Miocene)

Uzbekistan: Kara-tepe (Holocene)

See also Testudinidae