Abundance: 0.03 to 44.00% (median 0.95%)

Latitudinal range: -38.6° to 56.3°

Habitats: (43) only

Found in 43 samples

Argentina: Paso Otero 4 (water well) (Holocene)

Australia: Wee Jasper 99 (Pleistocene - Holocene), Mammoth Cave (Late Pleistocene), Beehive Cave (Holocene)

Canada: Charlie Lake Cave (Zone IV) (Holocene)

China: Shuangta (Holocene)

Ethiopia: Andalee (Middle Pleistocene)

Germany: Echzell (Early Miocene)

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E) (Late Pleistocene)

Lesotho: Sehonghong (GAP) (Holocene), Sehonghong (GWA) (Holocene), Sehonghong (ALP) (Holocene), Sehonghong (SA) (Holocene), Sehonghong (BARF) (Late Pleistocene), Sehonghong (RF) (Late Pleistocene), Sehonghong (OS) (Late Pleistocene), Sehonghong (MOS) (Late Pleistocene), Sehonghong (RFS) (Late Pleistocene)

Mexico: El Zurdo (Holocene), Santa Marta rock-shelter (Holocene)

Mongolia: Otson Tsokhio and Zuun Shovkh (Late Pleistocene), Zaraa Uul (Locus 2) (Holocene)

Namibia: Arrisdrift (Middle Miocene)

Niger: Garumele (context 1) (Holocene)

Russia: Muhkai II (layer 80) (Early Pleistocene), Zhukovskoe (Bed 6) (Late Pliocene)

Senegal: Diouboye (Holocene)

Spain: Can Roqueta II (Holocene)

Sri Lanka: Fa-Hien Lena Cave (Phase A) (Holocene), Fa-Hien Lena Cave (Phase C) (Late Pleistocene), Fa-Hien Lena Cave (Phase D) (Late Pleistocene)

Thailand: Spirit Cave (Layer 2) (Holocene), Spirit Cave (Layer 4) (Late Pleistocene)

Turkey: Kargi 1 (Late Oligocene), Bagiçi (Late Oligocene), Harami 1 (Late Oligocene)

United States: Vess Cave (Late Pleistocene), Steck (Holocene), Watson Brake (3.2 mm fraction) (Holocene), Guard (Holocene), Pine Hill (Holocene), Baker Village (Holocene)

Vietnam: Hiem Cave (Holocene)

See also Anura indet., Anura indet./"a number of bone fragments"