Abundance: 0.20 to 37.77% (median 9.83%)

Latitudinal range: 9.3° to 65.0°

Habitats: boreal forest/taiga (3), temperate broadleaf/mixed forest (9), temperate coniferous forest (6), temperate savanna (3), tropical/subtropical dry broadleaf forest (1), tropical/subtropical savanna (1)

Found in 23 samples

Austria: Thaya Valley

Cameroon: Garoua (1995)

Canada: Misery Bay Provincial Nature Reserve, Hat Creek Fire (mature forest)

China: Changbaishan Nature Reserve (mixed coniferous and broad-leaved forest), Wolong Natural Reserve (forest interior), Wolong Natural Reserve (forest edge), Wolong Natural Reserve (grassland)

Croatia: Mount Medvednica plot 1

Czech Republic: Ceske Budejovice

Finland: Nuuksio National Park (small traps, prop-glyc)

Germany: Ossweil (penncress-corn crop rotation)

Hungary: North Hungarian Mountain Range (control plots)

Italy: Natural Special Reserve of Fondo Toce (riparian wood)

Japan: Kitadan Valley

Russia: Indola River (biotope 1), Lazarevo (original forest area)

South Korea: Naju City (native forest), Naju City (managed forest), Naju City (forest patch)

United States: Coconino National Forest (1998), East Pond (wet meadow), Delta Junction

See also Carabus arcensis, Carabus auronitens, Carabus billbergi, Carabus canaliculatus, Carabus cancellatus, Carabus chamissonis, Carabus convexus, Carabus coriaceus, Carabus dehaanii, Carabus fraterculus, Carabus gigas, Carabus glabratus, Carabus granulatus, Carabus hortensis, Carabus hummeli, Carabus intricatus, Carabus jankowskii, Carabus maiyasanus, Carabus nemoralis, Carabus praecellens, Carabus scheidleri, Carabus seishinensis, Carabus serratus, Carabus shamaevi, Carabus sp., Carabus sylvosus, Carabus taedatus, Carabus ulrichii, Carabus vietinghoffi, Carabus vietinghoffii, Carabus violaceus