Trlica (Bed 10)
Basic information
Sample name: Trlica (Bed 10)

Reference: A. K. Agadzhanyan, I. A. Vislobokova, M. V. Shunkov, and V. A. Ulyanov. 2017. Pleistocene mammal fauna of the Trlica locality, Montenegro. Fossil Imprint 73(1-2):93-114 [ER 3942]
Geography
Country: Montenegro



Coordinate: 43° 20' 38" N, 19° 23' 0" E
Coordinate basis: stated in text

Time interval: Early Pleistocene

Unit number: 10

Unit order: above to below

Max Ma: 1.8

Min Ma: 1.5

Age basis: zone

Geography comments: "2.5 km southeast of the town of Pljevlja and 1.6 km northeast of the mouth of the Cehotina River canyon"
a lengthy discussion of biochronology undergirds the date of 1.8 to 1.5 Ma given in the abstract

Environment
Lithology: sandstone

Taphonomic context: carnivore accumulation,cave,rodent accumulation

Habitat comments: a "closed karstic cavity filled with loose deposits" and this bed is specifically "interbedding yellowish brown loams and loamy sands, travertine horizons, with interbeds rich in coprolite inclusions, teeth, and bones of large mammals and well traced scree paving. Bone of small mammals are relatively common... The bed is up to 60 cm thick" and was "formed mostly as proluvium"
the coprolites are "of hyenas... the cave was inhabited, periodically visited at least by hyenas and porcupines, and bats dwelt under the vault"
human artefacts are not mentioned and presumably absent

Methods
Life forms: bats,carnivores,rodents,ungulates,other small mammals

Sampling methods: quarry,screenwash

Sample size: 631 specimens

Years: 2010 - 2014

Sampling comments: "all sediments taken out of the cave were washed in sieves with 1 x 1 mm mesh"

Metadata
Sample number: 4345

Contributor: John Alroy

Enterer: John Alroy

Created: 2023-08-18 13:39:10

Modified: 2023-08-18 03:39:10

Abundance distribution
32 species
7 singletons
total count 631
geometric series index: 49.3
Fisher's α: 7.117
geometric series k: 0.8609
Hurlbert's PIE: 0.9000
Shannon's H: 2.6345
Good's u: 0.9889
Each square represents a species. Square sizes are proportional to counts.
Register
Ursus cf. etruscus3
plus 3 carnivores, 3 Ursus, and 5 Ursidae
Ursus deningeri32
Pachycrocuta brevirostris19
plus 2 Hyaenidae
Panthera cf. gombaszoegensis4
"Panthera onca cf. gombaszogensis"
Homotherium latidens10
"Homotherium crenatidens": see Anton et al. (2014)
Megantereon cultridens1
plus 2 Felidae
Canis etruscus8
plus 1 Canis
Canis lycanoides2
"Lycaon lycaonoides": plus 3 Vulpes of Nyctereutes
Mustelidae indet.3
Mammuthus ? meridionalis8
"Archidiskodon ? meridionalis": plus 8 Elephantidae, one possibly Palaeoloxodon
Palaeoloxodon sp.2
questionably referred
Stephanorhinus etruscus104
plus 1 Rhinocerotidae
Equus stenonis28
plus 4 Equus
Equus cf. major15
Praemegaceros sp.87
Cervus elaphus acoronatus60142 kg browser-grazer
"Cervus acoronatus": plus 13 Cervus
Capreolus sp.11
Cervalces gallicus1
"Libralces cf. gallicus": see Breda and Marchetti (2005); plus 2 Libralces and 3 Alcini
Bison (Eobison) sp.98
Leptobos cf. etruscus29
Gazellospira sp.1
Capra sp.5
questionably referred: plus 12 Caprinae
Megalovis balcanicus65
plus 4 possible Megalovis
Soergelia intermedia13
plus 5 possible Soergelia and 2 Bovidae
Rhinolophus macrorhinus2
compared to anomalidens
Myotis blythii longocaninus124.6 g
plus 5 Chiroptera
Mimomys pliocaenicus5
plus 19 in this species group
Mimomys reidi2
plus 11 in this group and 7 MImomys
Allophaiomys pliocaenicus1
plus 9 Microtini
Lagurus sp.1
Hystrix refossa9
plus 5 Hystrix and 51 Rodentia
Pliolagus sp.1
plus 4 Leporini